Cross-linking of DNA and protein was performed by treating the worms with 11% formaldehyde at 4°C for one hour. The worms were then lysed via Teflon homogenizer and sonicated to obtain 500-1000 base pair DNA fragments. ATFS-1 was then immunopreciptated from 500 mg of lysate using ATFS-1 antibodies [PMID 22700657] and protein A sepharose beads (Invitrogen). The cross-links were reversed by incubation at 65°C for 5 hours. The DNA fragments were then ethanol-precipitated and purified using QiaQuick Spin Columns (Qiagen). The DNA fragments were sequenced using the Hi-Seq Illumina platform.